Higher Classification: > Kingdom Animalia > Subkingdom Eumetazoa > Phylum Annelida > Class Polychaeta > Subclass Errantia > Order Phyllodocida > Suborder Aphroditiformia > Family Polynoidae > Subfamily Polynoinae > Genus Harmothoe

Harmothoe imbricata (Linnaeus, 1767)

Rank: Species
Taxon Status: accepted

Relationships towards this taxon

Genus group names

*Harmothoe*	Kinberg, 1856	accepted genus name
*Evarne*	Malmgren, 1865	junior homonym
*Harmothoe (Harmothoe)*	Kinberg, 1856	synonym
*Laenilla*	Malmgren, 1865	synonym
*Lagisca taurica*	(Czerniavsky, 1882)	synonym

Species group names

*Aphrodita lepidota*	Pallas, 1766	subjective synonym
*Aphrodita lepidota minuta*	Pennant, 1777	subjective synonym
*Aphrodita varians*	Dalyell, 1853	subjective synonym
*Aphrodita imbricata*	Linnaeus, 1767	synonym
*Harmothoe imbricata incerta*	(Bobretzky, 1881)	synonym
*Harmothoe maxillospinosa*	de Saint-Joseph, 1888	synonym

Vernaculars (+) (-)

Dutch: gladschubbige zeerups sources: Leewis, R. (2002). Flora en fauna van de zee [Marine flora and fauna]. Veldgids, 16. KNNV Uitgeverij: Utrecht, The Netherlands. ISBN 90-5011-153-X. 320 pp.

English: common fifteen-scaled worm; Fifteen-scaled worm; Free-living scale worm sources: Cole, T. C. H. (2017). Wörterbuch der Wirbellosen / Dictionary of Invertebrates. Latein-Deutsch-English. 2017. Springer Spektrum: Berlin. ISBN 978-3-662-52869-3. XII, 574 pp.;Volvenko, I. V.; Orlov, A. M.; Gebruk, A. V.; Katugin, O. N.; Ogorodnikova, A. A.; Vinogradov, G. M.; Maznikova, O. A. (2019). Checklist of the trawl macrofauna from the North Pacific and adjacent seas with information about fishery importance, potential product yield, and price range, supplement to: Volvenko, Igor V; Orlov, Alexei M; Gebruk, Andrey V; Katugin, Oleg N; Ogorodnikova, Alla A; Vinogradov, Georgy M; Maznikova, Olga A (in prep.): Trawl macrofauna of the Far-Eastern Seas and North Pacific: proportion of commercial species, potential product yield, and price rang. Earth System Science Data. PANGAEA - Data Publisher for Earth & Environmental Science.

German: gemeiner Schuppenwurm sources: Unterwasser-Welt Ostsee

Japanese: マダラウロコムシ sources: Japan Agency for Marine-Earth Science and Technology. (2009 onwards). Biological Information System for Marine Life (BISMaL). Accessed on 2018-11-21.

Russian: Хармотое черепитчатая

Literature

additional source	Pettibone, Marian H. (1963). Marine polychaete worms of the New England region. I. Aphroditidae through Trochochaetidae. Bulletin of the United States National Museum. 227(1): 1-356.
additional source	Malmgren, Anders Johan. (1865). Nordiska Hafs-Annulater. [part one of three]. Öfversigt af Königlich Vetenskapsakademiens förhandlingar, Stockholm. 22(1): 51-110, plates VIII-XV.
additional source	Hartman, Olga. (1938). The types of the polychaete worms of the families Polynoidae and Polyodontidae in the United States National Museum and the description of a new genus. Proceedings of the United States National Museum. 86(3046): 107-134.
additional source	Trott, T. J. (2004). Cobscook Bay inventory: a historical checklist of marine invertebrates spanning 162 years. Northeastern Naturalist. 11, 261-324.
additional source	Hartmann-Schröder, G. (1996). Annelida, Borstenwürmer, Polychaeta [Annelida, bristleworms, Polychaeta]. 2nd revised ed. The fauna of Germany and adjacent seas with their characteristics and ecology, 58. Gustav Fischer: Jena, Germany. ISBN 3-437-35038-2. 648 pp.
additional source	Pollock, L.W. (1998). A practical guide to the marine animals of northeastern North America. Rutgers University Press. New Brunswick, New Jersey & London. 367 pp.
additional source	Brunel, P., L. Bosse & G. Lamarche. (1998). Catalogue of the marine invertebrates of the estuary and Gulf of St. Lawrence. Canadian Special Publication of Fisheries and Aquatic Sciences, 126. 405 pp.
additional source	Johnston, G. (1865). A catalogue of the British non-parasitical worms in the collection of the British Museum. [book]. 1-365. British Museum. London. [See also separate entry for Baird supplement].
additional source	David, Andrew A.; Cahill, Jordan. (2020). Tri-oceanic connectivity of the supposedly cosmopolitan polychaete, Harmothoe imbricata (Annelida: Polynoidae): insights from the COI marker. Marine Biology Research. 16(4): 256-264.
additional source	Nygren, Arne; Norlinder, Erika; Panova, Marina ; Pleijel, Fredrik. (2011). Colour polymorphism in the polychaete Harmothoe imbricata (Linnaeus, 1767). Marine Biology Research. 7(1): 54 - 62.
additional source	Malmgren, A.J. (1867). Annulata Polychaeta Spetsbergiæ, Grœnlandiæ, Islandiæ et Scandinaviæ. Hactenus Cognita. Ex Officina Frenckelliana, Helsingforslæ. 127 pp. & XIV plates.
additional source	Hartman, Olga. (1959). Catalogue of the Polychaetous Annelids of the World. Parts 1 and 2. Allan Hancock Foundation Occasional Paper. 23: 1-628.
additional source	Fishelson, L.; Rullier, F. (1969). The Second South Red Sea Expedition, 1965, Report No. 9. Quelques Annélides Polychètes de la Mer Rouge. Israel Journal of Zoology. 18: 49-117.
additional source	McIntosh, W.C. 1900. A monograph of British Annelids. vol.1. pt.2. Polychaeta Amphinomidae to Sigalionidae. Ray Society of London, 1(l2): 215-442.
additional source	Liu, J.Y. [Ruiyu] (ed.). (2008). Checklist of marine biota of China seas. China Science Press. 1267 pp.
additional source	Johnson, Herbert P. 1897. A preliminary account of the marine annelids of the Pacific coast, with descriptions of new species. Proceedings of the California Academy of Sciences, Ser. 3, 1(5): 153-199.
source of synonymy	Imajima, Minoru. (1997). Polychaetous annelids from Sagami Bay and Sagami Sea collected by the Emperor Showa of Japan and deposited at the Showa Memorial Institute, National Science Museum, Tokyo. Families Polynoidae and Acoetidae. National Science Museum Monographs. 13: 1-131.
redescription	Ruff, R. Eugene. (1995). Family Polynoidae Malmgren, 1867. 105-166. IN: Blake, James A., Hilbig, Brigitte, and Scott, Paul H. (Ed.). Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Volume 5 - The Annelida Part 2. Polychaeta: Phyllodocida (Syllidae and scale-bearing families), Amphinomida, and Eunicida. Santa Barbara Museum of Natural History. Santa Barbara.
redescription	Jirkov, I.A. (2001). [Polychaeta of the Arctic Ocean] (In Russian) Polikhety severnogo Ledovitogo Okeana. Yanus-K Press, Moscow, 632 pp.
redescription	Imajima, Minoru. (1997). Polychaetous annelids from Sagami Bay and Sagami Sea collected by the Emperor Showa of Japan and deposited at the Showa Memorial Institute, National Science Museum, Tokyo. Families Polynoidae and Acoetidae. National Science Museum Monographs. 13: 1-131.
identification resource	Tebble, Norman; Chambers, Susan. (1982). Polychaetes from Scottish Waters, Part 1. Family Polynoidae. Royal Scottish Museum Studies. [unnumbered] : 1-73.
biology source	Garm, Anders; Simonsen, Sidsel H.; Mendoza-González, Paula; Worsaae, Katrine; ;. (2021). Have the eyes of bioluminescent scale worms adapted to see their own light? A comparative study of eyes and vision in Harmothoe imbricata and Lepidonotus squamatus. Journal of Experimental Biology. 224(14): ?-?.

Genbank

Citation

Read, G.; Fauchald, K. (Ed.) (2025). World Polychaeta Database. Harmothoe imbricata (Linnaeus, 1767). Accessed through: World Register of Marine Species at: https://marinespecies.org/aphia.php?p=taxdetails&id=130769